What a good question! I never realised it was so difficult. My immediate answer was the same as LWAB's first response, above.
If we start by supposing the circumference of an 18T sprocket is 9" and that of a 19T sprocket is 9.5" (in practice this is a bit too much, what we need is the circumference around the chain rivets that is approximately halfway down the teeth of the sprocket, but I'll assume the error is negligible). It follows the difference in radius between 18T and 19T is (9.5 - 9)/2/PI or 1/4/PI = 0.08"
So I
think the answer is that the wheel moves forward by the difference in radius of an 18T and 19T sprocket. The (fixed length) chain will just wrap to accommodate this difference.
Doesn't sound much does it? I'm not at all confident there isn't a mistake somewhere here. Perhaps Ben T should just do it, take measurements and let us know; making sure there's no change in chain tension of course