If I cycle over mixed terrain for a long time (a whole day out) I always find that I have averaged about 14mph.
Walking cross-country over mixed terrain I can sustain a speed of 3.25mph almost indefinitely.
Oh, and a 400k Audax used to take me 24 hours, and so does a 100k cross-country walk. So the 4:1 ratio seems pretty much right to me for fairly brisk
walking and its cycling equivalent, that is transportational, non-racing riding.
But whether the 4:1 ratio holds true for cycling at racing speeds versus running, I couldn't say. The foot-strike-induced fatigue of running (especially on hard surfaces) is much more significant than that of walking.
There's no significant impact-related fatigue in road cycling; but there
is fatigue associated with maintaining one's position on the bike.
Although, having said that, I ride recumbents, so it doesn't really apply to me